Malacostraca | 
Decapoda | 
Varunidae
			
			
			
				Environment: milieu / climate zone / depth range / distribution range				
				
					Ecology				
				
			
			
				Benthic; freshwater; brackish; depth range 0 - 25 m (Ref. 81172).  Temperate; 5°C - 27°C (Ref. 8003); 44°N -   21°N, 112°E -   135°E			
			
			
				
			
			
			
				Northwest Pacific:  Native from Valdivostok Russia, to the west coast of North Korea south to Hong Kong, China.  Introduced to central and western Europe.
			
			
			
			
			
				Length at first maturity / Size / Weight / Age
			
			
				Maturity: Lm ?  range ? - ? cm Max length : 7.5 cm CW male/unsexed; (Ref. 3159)			
			
			
						
			
			
			
			
				
					Catadromous (Ref. 101972).  Maximum depth from Ref. 104445. Euryhaline (Ref. 115548).  Catadromous (Ref. 115520, 115522), spending most of its life growing to maturity in freshwater (Ref. 115522).  Found in coastal rivers and estuaries.  It needs brackish or saltwater to reproduce (Ref. 8778).  Downstream migration of adults occurs worldwide (Ref. 115548).  From sublittoral sediment and surface running water (Ref. 104191).  Adults are found in fresh, brackish and salt waters; oviparous females are typically in greatest number in saltwater (Ref. 115548).  Has high affinity to burrow.  Juveniles are known to form burrows in soft-sediment banks between high and low tide lines upon  migration into brackish channels and creeks.  In general, small juveniles construct and inhabit these burrows (Ref. 104191).  Larger crabs occasionally burrow in softer substrate of freshwater stream banks.  When abundance increases, burrows can become very dense, causing bank weakening, erosion, loss of bank vegetation, and bank collapse (Ref. 115547).  In Germany, burrows reached 50 cm deep, which caused considerable damage (Ref. 115521).  Predominantly omnivorous; however, it exhibits plasticity in its diet (i.e., feeding habits may shift throughout the cycle).  Likely to scavenge and feed on detritus (Ref. 104191).  Larvae feed on phytoplankton; newly settled juveniles feed on aquatic plants (Ref. 115548).  Shifts to a more carnivorous diet as it ages, eating shrimp and other benthic invertebrates.  Also reported to feed on fish captured in fishing nets (Ref. 104191).  Opportunistic feeder (Ref. 115548).  Feeds on periphytic algae, terrestrial plants, detritus, sand, copepods and chironomid larvae (Ref. 107018).  Carries a lung fluke trematode parasite Paragonimus westermani that infects humans (Ref. 115522).				
			
			
			
			
			
			
				
					Life cycle:  Eggs are carried until hatched and the adults die afterwards.  The larvae undergo five zoeal stages.  Juveniles migrate from the estuary to the freshwater to develop into adults.  Adults migrate downstream, mating occurs in saline water.				
			
			
			
			
			
				Ray, G.L. 2005 Invasive Animal Species in Marine and Estuarine Environments: Biology and Ecology. Aquatic Nuisance Species Research Program. (Ref. 3159)
			
			
			
			
			
			
			
					
						IUCN Red List Status    
						 (Ref. 130435: Version 2025-1)
					
					
					
				
			
			
			
			
				CITES status   (Ref. 108899)
			
			
				
					Not Evaluated				
			
			
			
			
			
			
				
					Not Evaluated				
			
			
			
			
			
				Threat to humans  
			
			
				
									
			
			
			
			
			
			
			
			
				Human uses  
			
			
				Fisheries: commercial			
			
			
				FAO - Aquaculture: production, species profile; Fisheries: landings, species profile | FishSource | Sea Around Us			
			
			
			
			
			
Tools
			
			
			
			
				
					More information				
				
					 PhysiologyOxygen consumption
  Human RelatedStamps, coins, misc.
  
			 
			
			
			
				Internet sources
			
			
			
			
			
				Estimates based on models
			
			
			
									
						Preferred temperature  					
				 (Ref. 
115969): 14.1 - 23.2, mean 20 (based on 109 cells).			
 
			
			
			
			
			
						
							
				
					
						Fishing Vulnerability  					
					
					
						Low to moderate vulnerability (29 of 100).					
				
						
						
			
									
						Price category  					
					
					Unknown.