| Citation | Bisby, F.A., Y.R. Roskov, T.M. Orrell, D. Nicolson, L.E. Paglinawan, N. Bailly, P.M. Kirk, T. Bourgoin, G. Baillargeon and D. Ouvrard 2011 Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist. Digital resource at www.catalogueoflife.org/annual-checklist/2011/. Species 2000: Reading, UK. | 
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| Ref. No. | 88916 | 
| Language | English | 
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