| شماره مرجع | Brown, W.Y., 1976 |
| بوم سازگان _ اكوسيستم | Hawaii |
| وضعیت | native |
| فراواني | |
| Lifestage | adults |
| Regional database | |
| Remarks | Annual and regular breeding cycle; peak egg-laying: Manana: last week of March or the first week of April; Kure: May; Johnston Atoll: middle of March (Ref. 96995). Also in Ref. 84934. C: Refs. 80078, 84934; O: Refs. 80078, 96995. |